The mandible (from Latin: mandibula or mandĭbŭ-lum, a jaw) [1] of an arthropod is a pair of mouthparts used for either for biting, cutting and holding food. The last three cephalic segments, together with the three most anterior thoracic segments (all of the cephalothorax), house the external mouthparts. From anterior to posterior these are the mandibles, maxillules, maxillae and then the three pairs of thoracic maxillipeds. These are all biramous except for the mandibles and maxillules. [2] Mandibles are often simply referred to as jaws. Maxillipeds are appendages modified to function as mouthparts.  Hermit crabs are often seen grooming their eyes with their maxillipeds much like a cat uses it’s paw to clean its face.

Coenobita mandibles and maxipelllids Credit – Chen Yu Jung
Cavipes mouth
Cavipes mouth
Coenobita mouth mandibles maxillipeds
Rugosus mouth
Rugosus mouth
Coenobita mouth mandibles maxillipeds
Coenobita mandibles and maxipelllids Credit – Chen Yu Jung
Coenobita mouth mandibles maxillipeds
Coenobita mandibles and maxipelllids Credit – Chen Yu Jung
Coenobita mouth mandibles maxillipeds
C. perlatus Photo Credit – Mary Milhorn
Coenobita male – Photo credit – Mary Milhorn

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Overview of the anatomy of a land hermit crab (Coenobita)

Photo Credits:
The Crab Street Journal has been granted permission by these photographers to use their photo(s) on our site.
Chen Yu-Jung

References:
1 Latin Dictionary Founded on Andrews’ edition of Freund’s Latin dictionary revised by Charlton T. Lewis, Ph.D. and. Charles Short, LL.D. Oxford. Clarendon Press. 1879.

2. Dardanus megistos by Storm Martin 2012